UTP18


This analysis report summarizes UTP18 eCLIP data from ENCODE. The current tab provides an overview of the results, and the other tabs list identified peaks and motifs.

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Experiment Summary


Experiment ID File ID Sample Antibody source Antibody ID Antibody lot Release date Laboratory Number of peaks
ENCSR488JKQENCFF644EIRHepG2Bethyl LabsA301-551A12018-08-30Gene Yeo, UCSD47
ENCSR964VOXENCFF783INOK562Bethyl LabsA301-551A12018-08-30Gene Yeo, UCSD133

UTP18 Peak Assignments


The following plots show the assignment of peaks into different genic features:

Introns contain other subdivisions, please see the subsequent plot for a breakdown of those features.

This plot is interactive. You can hover over the bars to see the percentage and number of identified peaks. You can select which features to show by toggling the region names in the legend, and you can download the plot by moving your mouse to the top of the plot and clicking on the camera icon.

UTP18 Intron Assignments


The following plots show the assignment of peaks into different intronic features:

  •  5’ splice site: the peak is within the first 100 bp of an intron (5’ to 3’ direction
  •  3’ splice site: the peak is within the last 100 bp of an intron (5’ to 3’ direction)
  •  Proximal intron: the peak is within 100 bp to 500 bp of the nearest exon
  •  Distal intron: greater than 500 bp away from the nearest exon

This plot is interactive. You can hover over the bars to see the percentage and number of identified peaks. You can select which features to show by toggling the region names in the legend, and you can download the plot by moving your mouse to the top of the plot and clicking on the camera icon.

UTP18 Motifs


The following plot shows a summary of the top five motifs by significance in each experiment. Motifs were detected with varying length parameters, you may observe a similar motif with different number of bases included multiple times. A motif present in more peaks and with a higher significance will be in the upper right quadrant of the plot, while the lower left quadrant has motifs that are less significant and in fewer peaks.

This plot is interactive. You can hover over the bars to see the percentage and number of identified peaks. You can select which features to show by toggling the region names in the legend, and you can download the plot by moving your mouse to the top of the plot and clicking on the camera icon.



The following tables contain called UTP18 peaks. To identify peaks, clusters (regions of read enrichment) in the immunoprecipitated (IP) samples were found using the peak calling tool CLIPper. To account for background signal, a cluster was identified as a peak if the log2 fold enrichment over input was ≥ 3 and the p-value ≤ 0.001.

Peaks were annotated using transcript information from GENCODE. Each annotated peak is labeled with specific annotation feature types, first split by coding and non-coding transcripts, then by transcript regions, and then by intron/exon proximity regions. For overlapping transcript regions, the following hierarchy is used to label the region: coding sequence (CDS), 5’ or 3’ untranslated region (UTR), intron, non-coding exon, then non-coding intron. For example, if a peak is in the 5’ UTR of one transcript that overlaps an intron of another transcript, the peak region will be labeled as 5’ UTR. All gene annotations are from GENCODE release v41.

The following columns are included in the tables:

  •  Chromosome: chromosome where the peak is located.
  •  Peak start: start coordinate for the peak.
  •  Peak stop: stop coordinate for the peak.
  •  Strand: strand of DNA where the peak was called.
  •  -log10(p-value): significance of the cluster call by CLIPper. Values are log transformed so a larger number indicates a greater significance.
  •  log2(fold change): log transformed fold enrichment of the IP signal over a matched input.
  •  Gene name: name (symbol) of the gene that overlaps with the peak.
  •  Ensembl ID: unique identifer of the gene that overlaps with the peak.
  •  Feature: which part of the listed gene overlaps with the peak.

Please note, this table is interactive. You can search for specific genes or features, and can sort the table by any of the columns.

ENCSR488JKQ (HepG2) Peaks


Chromosome Peak start Peak stop Strand -log10(p-value) log2(fold change) Gene name Ensembl ID Feature
chr1117,075,80517,075,823-400.0003.326SNORD14BENSG00000201403.1ncRNA
chr2026,209,31826,209,338-75.8516.257MIR663AHGENSG00000227195.11ncRNA
chr2026,209,33826,209,386-79.7636.297MIR663AHGENSG00000227195.11ncRNA
chr218,211,5158,211,560+8.7213.858ENSG00000278996ENSG00000278996.1ncRNA
chr9114,311,653114,311,722+13.3273.176COL27A1ENSG00000196739.153' UTR
chr1719,190,14119,190,177-400.0007.272SNORD3CENSG00000264940.5ncRNA
chr1570,097,64370,097,778-5.9643.819TLE3ENSG00000140332.185' UTR
chr898,825,40898,825,441-9.2966.181STK3ENSG00000104375.175' splice site
chr2063,944,11763,944,196-9.9333.044UCKL1ENSG00000198276.16Proximal intron
chr162,530,6882,530,800+8.4793.417AMDHD2ENSG00000162066.16CDS
chr161,965,2651,965,292+8.4793.417SNORA78|SNHG9ENSG00000273587.1||ENSG00000255198.6ncRNA
chr1719,190,17719,190,213-400.0006.833SNORD3CENSG00000264940.5ncRNA
chr9114,240,216114,240,266+9.7813.197COL27A1ENSG00000196739.15CDS
chr1719,062,06119,062,119+170.24110.249SNORD3B-1ENSG00000265185.6ncRNA
chr1719,062,11919,062,140+154.11010.078SNORD3B-1ENSG00000265185.6ncRNA
chr9114,252,592114,252,628+7.8753.378COL27A1ENSG00000196739.15CDS
chr2231,463,199231,463,238-20.2893.332NCLENSG00000115053.17CDS
chr2231,463,238231,463,299-13.1573.026NCLENSG00000115053.17CDS
chr773,593,36673,593,393-5.6943.584MLXIPLENSG00000009950.163' UTR
chr1719,190,10619,190,141-400.0006.850SNORD3CENSG00000264940.5ncRNA
chr14105,468,639105,468,687+5.2004.263MTA1ENSG00000182979.18Proximal intron
chr9114,231,822114,231,848+4.7313.070COL27A1ENSG00000196739.15CDS
chr1117,075,84217,075,865-400.0004.333SNORD14BENSG00000201403.1ncRNA
chr1117,075,82317,075,842-400.0003.667SNORD14BENSG00000201403.1ncRNA
chr142,943,23442,943,259-4.7583.123SLC2A1ENSG00000117394.24CDS
chr339,411,11539,411,165+16.1853.687SNORA62ENSG00000202363.1ncRNA
chr218,401,8188,401,848+119.56010.002ENSG00000280441ENSG00000280441.3ncRNA
chr218,401,7838,401,818+400.0007.549ENSG00000280441ENSG00000280441.3ncRNA
chr218,401,7798,401,783+27.2787.028ENSG00000280441ENSG00000280441.3ncRNA
chr218,218,7408,218,832+400.0003.380ENSG00000278996ENSG00000278996.1ncRNA
chr2017,629,84917,629,943-10.8943.033RRBP1ENSG00000125844.17CDS
chr2231,460,475231,460,567-23.1013.310NCLENSG00000115053.17CDS
chr2231,461,978231,462,018-12.7703.008NCLENSG00000115053.17CDS
chr626,055,77726,055,816-12.0473.490H1-2ENSG00000187837.4CDS
chr9114,235,636114,235,651+5.9643.876COL27A1ENSG00000196739.15CDS
chr1719,188,09919,188,107+400.0006.206SNORD3AENSG00000263934.5ncRNA
chr1719,188,10719,188,131+400.0006.173SNORD3AENSG00000263934.5ncRNA
chr1719,188,09419,188,099+400.0006.141SNORD3AENSG00000263934.5ncRNA
chr1719,188,06819,188,094+400.0005.977SNORD3AENSG00000263934.5ncRNA
chr1176,883,39776,883,411+26.9246.717ACER3ENSG00000078124.13Distal intron
chr1176,883,41176,883,419+26.9246.717ACER3ENSG00000078124.13Distal intron
chr2231,461,593231,461,678-12.8413.226NCLENSG00000115053.17CDS
chr1719,112,55219,112,617-63.8987.249ENSG00000262202|SNORD3DENSG00000262202.4||ENSG00000277947.1||ENSG00000281000.1ncRNA
chr580,651,22880,651,311-13.9533.497DHFRENSG00000228716.7Distal intron
chr72,354,9132,354,971+5.6293.756EIF3BENSG00000106263.19CDS
chr202,839,0212,839,084-6.0873.141PCED1AENSG00000132635.17CDS
chr9114,237,687114,237,715+6.2393.330COL27A1ENSG00000196739.15CDS


ENCSR964VOX (K562) Peaks


Chromosome Peak start Peak stop Strand -log10(p-value) log2(fold change) Gene name Ensembl ID Feature
chr2202,293,075202,293,103+6.2714.301SNORD11ENSG00000238317.2ncRNA
chr218,209,1488,209,226+17.4426.746hsa-mir-3648-1MI0016048miRNA
chr1162,854,63062,854,651-26.2583.401SNORD28|SNHG1ENSG00000274544.1||ENSG00000255717.9ncRNA
chrX153,691,264153,691,299+4.8713.342SLC6A8ENSG00000130821.173' splice site
chr452,712,70352,712,739+14.4323.374hsa-mir-4449MI0016792miRNA
chr1228,146,081228,146,156+5.5704.445GUK1ENSG00000143774.175' splice site
chr1782,097,16582,097,264-19.9603.680FASNENSG00000169710.95' UTR
chr11123,058,119123,058,164-85.7323.102SNORD14EENSG00000200879.1ncRNA
chr3160,514,940160,515,003-72.3214.041SCARNA7ENSG00000238741.1ncRNA
chr1205,749,959205,750,023-15.6403.204NUCKS1ENSG00000069275.13CDS
chr7148,883,795148,883,895-6.9243.623EZH2ENSG00000106462.12Proximal intron
chr1689,588,43889,588,558+3.8123.021CPNE7ENSG00000178773.15Proximal intron
chr998,015,42298,015,480+25.8194.170ANP32BENSG00000136938.9CDS
chr218,210,7168,210,804+27.3437.790ENSG00000278996ENSG00000278996.1ncRNA
chr9136,668,092136,668,148+5.7383.927EGFL7ENSG00000172889.163' splice site
chr227,383,95127,384,017-4.1573.560PPM1GENSG00000115241.11CDS
chr75,529,8245,529,880-12.3223.057ACTBENSG00000075624.175' UTR
chr1173,866,877173,866,890-69.3074.718GAS5ENSG00000234741.10ncRNA
chr1173,866,890173,866,927-84.9174.460GAS5ENSG00000234741.10ncRNA
chr1173,866,927173,866,939-13.0833.623GAS5ENSG00000234741.10ncRNA
chr71,453,2471,453,311-5.5845.306MICALL2ENSG00000164877.19Distal intron
chr218,401,8188,401,851+400.00010.317ENSG00000280441ENSG00000280441.3ncRNA
chr3128,492,950128,493,033+4.5523.556GATA2-AS1ENSG00000244300.3ncRNA
chr1949,497,73249,497,791+29.0023.418SNORD35BENSG00000200530.1ncRNA
chr1117,075,82517,075,867-44.3993.817SNORD14BENSG00000201403.1ncRNA
chr1566,502,02166,502,082-20.1143.226ENSG00000280554|SNORD18BENSG00000202529.1||ENSG00000280554.1ncRNA
chr161,376,2571,376,305-3.5674.945UNKLENSG00000059145.19Distal intron
chr218,397,6538,397,673+22.7735.313ENSG00000280441ENSG00000280441.3ncRNA
chr218,397,6208,397,653+11.3893.865ENSG00000280441ENSG00000280441.3ncRNA
chr1912,703,59912,703,671-18.4493.474hsa-mir-10395MI0033419miRNA
chr1117,075,80917,075,820-57.9204.200SNORD14BENSG00000201403.1ncRNA
chr1117,075,77617,075,809-39.2173.929SNORD14BENSG00000201403.1ncRNA
chr175,787,91475,787,953+51.6823.282SNORD45AENSG00000207241.1ncRNA
chr41,245,3461,245,437-5.5845.369CTBP1ENSG00000159692.17Distal intron
chr218,401,7828,401,789+304.52310.897ENSG00000280441ENSG00000280441.3ncRNA
chr218,401,7898,401,818+400.00010.472ENSG00000280441ENSG00000280441.3ncRNA
chr218,401,7808,401,782+111.4039.666ENSG00000280441ENSG00000280441.3ncRNA
chr1566,501,25766,501,315-6.8233.187SNORD18CENSG00000199574.1ncRNA
chr1776,561,64676,561,701+6.9674.125SNORD1A|SNHG16ENSG00000278261.1||ENSG00000163597.17ncRNA
chr218,398,3688,398,387+8.3255.895ENSG00000280441ENSG00000280441.3ncRNA
chr218,398,3608,398,368+8.3255.895ENSG00000280441ENSG00000280441.3ncRNA
chr1728,720,54928,720,618+38.3343.862RPL23AENSG00000198242.145' UTR
chr5138,561,038138,561,065-22.2394.026SNORD63ENSG00000206989.1ncRNA
chr5138,561,069138,561,080-20.3923.467SNORD63ENSG00000206989.1ncRNA
chr798,881,74698,881,805+70.5963.728hsa-mir-3609MI0015999miRNA
chr798,881,81698,881,880+71.4193.620hsa-mir-3609MI0015999miRNA
chr1776,561,10176,561,182+7.5364.471SNORD1BENSG00000199961.1ncRNA
chr11123,058,907123,058,913-16.6554.343HSPA8ENSG00000109971.145' UTR
chr11123,058,913123,058,952-31.6513.423HSPA8ENSG00000109971.145' UTR
chr191,421,7051,421,777+4.9254.427DAZAP1ENSG00000071626.17Proximal intron
chr1670,529,52470,529,590+5.4833.153SNORD111BENSG00000221514.1ncRNA
chr162,769,3072,769,354+8.2054.895SRRM2ENSG00000167978.175' splice site
chr218,218,7408,218,769+400.0007.567ENSG00000278996ENSG00000278996.1ncRNA
chr218,218,7698,218,781+400.0007.312ENSG00000278996ENSG00000278996.1ncRNA
chr218,218,7818,218,809+400.0006.770ENSG00000278996ENSG00000278996.1ncRNA
chr218,218,8098,218,887+400.0006.616ENSG00000278996ENSG00000278996.1ncRNA
chr1193,721,67693,721,726+46.1234.559SCARNA9ENSG00000254911.3ncRNA
chr126,510,5026,510,543+18.8864.308SCARNA10|ENSG00000276232ENSG00000239002.3||ENSG00000276232.1ncRNA
chr126,510,4516,510,495+44.5403.928ENSG00000276232|SCARNA10ENSG00000239002.3||ENSG00000276232.1ncRNA
chr2231,456,443231,456,525-22.6733.421SNORD20ENSG00000207280.1ncRNA
chr192,270,2322,270,279+6.2214.625OAZ1ENSG00000104904.12Proximal intron
chr2158,457,100158,457,220+5.6354.180PKP4ENSG00000144283.235' UTR
chr1728,723,69428,723,743+5.7384.003RPL23AENSG00000198242.14CDS
chr1193,721,75093,721,787+400.0004.111SCARNA9ENSG00000254911.3ncRNA
chr1391,348,18691,348,252+18.3033.784MIR17HGENSG00000215417.13ncRNA
chr933,952,77533,952,819-4.2335.056SNORD121AENSG00000238886.1ncRNA
chr218,218,4068,218,419+24.0377.201ENSG00000278996ENSG00000278996.1ncRNA
chr218,218,4198,218,441+25.4406.349ENSG00000278996ENSG00000278996.1ncRNA
chr2202,276,440202,276,509+12.7954.987SNORD70ENSG00000212534.1ncRNA
chr19893,560893,590+7.4983.719RNU6-9ENSG00000207507.1ncRNA
chr1719,190,10119,190,188-400.0006.663SNORD3CENSG00000264940.5ncRNA
chr3186,784,818186,784,846+25.3663.370SNORD2ENSG00000238942.1ncRNA
chr41,802,1111,802,194+5.0854.556FGFR3ENSG00000068078.205' splice site
chr126,769,9236,770,000+6.0153.172PTMSENSG00000159335.18CDS
chr2049,280,34449,280,383+47.6083.168SNORD12B|ZFAS1ENSG00000177410.13||ENSG00000222365.1ncRNA
chrX136,879,227136,879,275-28.1443.498SNORD61ENSG00000206979.1ncRNA
chr9133,350,099133,350,151+7.3233.265SNORD36BENSG00000200831.1ncRNA
chr1109,100,286109,100,299+400.0003.594ENSG00000270066|SCARNA2ENSG00000270066.3||ENSG00000278249.1ncRNA
chr1109,100,257109,100,275+400.0003.534ENSG00000270066|SCARNA2ENSG00000270066.3||ENSG00000278249.1ncRNA
chr1109,100,275109,100,285+400.0003.515SCARNA2|ENSG00000270066ENSG00000278249.1||ENSG00000270066.3ncRNA
chr9133,350,846133,350,930+55.9183.570SNORD36CENSG00000252542.1ncRNA
chr1109,100,299109,100,309+400.0003.615SCARNA2|ENSG00000270066ENSG00000278249.1||ENSG00000270066.3ncRNA
chr1719,112,54719,112,594-400.0005.275ENSG00000262202|SNORD3DENSG00000262202.4||ENSG00000281000.1||ENSG00000277947.1ncRNA
chr1068,755,18068,755,236+6.9674.395SNORD98ENSG00000283551.1ncRNA
chr1719,188,10419,188,128+400.0005.035SNORD3AENSG00000263934.5ncRNA
chr9136,726,771136,726,814-59.2143.095SNHG7|SNORA17A|ENSG00000281808ENSG00000274998.1||ENSG00000281808.1||ENSG00000233016.9ncRNA
chr9133,349,388133,349,440+23.8353.440SNORD24ENSG00000206611.1ncRNA
chr9133,349,440133,349,471+24.9923.409SNORD24ENSG00000206611.1ncRNA
chr115,254,3575,254,392-6.6123.521ENSG00000284931|HBG2ENSG00000284931.1||ENSG00000196565.15CDS
chr75,530,1725,530,252-12.7653.105ACTBENSG00000075624.17Proximal intron
chr1719,062,03519,062,137+249.2949.578SNORD3B-1ENSG00000265185.6ncRNA
chr172,329,1282,329,197-21.2223.729SNORD91BENSG00000275084.4ncRNA
chr2222,557,46322,557,544-26.0263.024PRAMEENSG00000185686.185' UTR
chr1193,721,82193,721,859+67.9355.400ENSG00000273885|SCARNA9ENSG00000273885.1||ENSG00000254911.3ncRNA
chr1109,100,309109,100,314+400.0003.304SCARNA2|ENSG00000270066ENSG00000278249.1||ENSG00000270066.3ncRNA
chr1109,100,315109,100,331+400.0003.046SCARNA2|ENSG00000270066ENSG00000270066.3||ENSG00000278249.1ncRNA
chr4173,331,981173,332,088-18.5713.072HMGB2ENSG00000164104.12CDS
chr126,472,69526,472,767+11.7943.799HMGN2ENSG00000198830.115' splice site
chr191,408,2771,408,368+9.1863.653DAZAP1ENSG00000071626.17Proximal intron
chr1175,400,39075,400,517+24.0353.086RPS3ENSG00000149273.155' UTR
chr2026,209,31626,209,369-4.9065.616MIR663AHGENSG00000227195.11ncRNA
chr1165,443,56565,443,632+7.7923.712NEAT1ENSG00000245532.11ncRNA
chr12132,550,461132,550,543+5.1385.162FBRSL1ENSG00000112787.14Distal intron
chr998,015,40098,015,421+16.3933.573ANP32BENSG00000136938.9CDS
chrX20,136,14120,136,192-8.6184.454EIF1AXENSG00000173674.11Proximal intron
chrX20,136,12020,136,141-6.8744.106ENSG00000201882ENSG00000201882.1ncRNA
chr193,982,5173,982,557-16.1343.795SNORD37ENSG00000206775.1ncRNA
chr1193,733,26093,733,294-4.7203.303hsa-mir-1304MI0006371miRNA
chr1173,866,317173,866,354-16.7254.369GAS5ENSG00000234741.10ncRNA
chr1193,721,57993,721,613+17.2623.240ENSG00000275146|SCARNA9ENSG00000275146.1||ENSG00000254911.3ncRNA
chr202,656,6442,656,681+32.8783.802SNORD56ENSG00000229686.1ncRNA
chr2202,291,373202,291,399+7.0713.784SNORD11BENSG00000271852.1ncRNA
chr482,895,85382,895,910-10.2215.512THAP9-AS1ENSG00000251022.6ncRNA
chr12103,947,672103,947,710+6.4723.924HSP90B1ENSG00000166598.163' UTR
chr172,330,2812,330,351-26.4364.086SNORD91AENSG00000212163.6ncRNA
chr1728,723,43728,723,488+6.9525.806SNORD42AENSG00000238649.1ncRNA
chr296,894,24696,894,298-5.1433.572FAM178BENSG00000168754.15Proximal intron
chr540,832,66240,832,735-8.3255.849SNORD72ENSG00000212296.1ncRNA
chr1688,966,43288,966,476-3.6594.188CBFA2T3ENSG00000129993.15Distal intron
chr10100,237,088100,237,143-13.6444.193CWF19L1ENSG00000095485.18Proximal intron
chr1719,112,59419,112,615-400.0005.045SNORD3D|ENSG00000262202ENSG00000262202.4||ENSG00000277947.1||ENSG00000281000.1ncRNA
chr130,968,16430,968,280-8.6184.120PUM1ENSG00000134644.165' splice site
chr128,578,75528,578,811-6.0363.228SNORD99ENSG00000221539.1ncRNA
chr1685,615,25885,615,317+4.9254.427GSE1ENSG00000131149.19Distal intron
chr11123,059,332123,059,378-400.0003.355HSPA8ENSG00000109971.145' UTR
chr218,218,0478,218,063+14.1696.631ENSG00000278996ENSG00000278996.1ncRNA
chr218,218,0168,218,039+31.1244.000ENSG00000278996ENSG00000278996.1ncRNA
chr218,217,9838,218,016+11.3103.134ENSG00000278996ENSG00000278996.1ncRNA
chr1117,074,69517,074,739-15.2694.850SNORD14AENSG00000272034.1ncRNA
chr1117,074,65117,074,695-29.0023.494SNORD14AENSG00000272034.1ncRNA
chr6132,815,330132,815,364+52.2774.091SNORD101ENSG00000206754.1ncRNA
chr1719,188,10019,188,104+400.0005.065SNORD3AENSG00000263934.5ncRNA
chr1719,188,06719,188,100+400.0004.889SNORD3AENSG00000263934.5ncRNA

The following tables contain identified motifs, where a motif is a short sequence of nucleotides that is significantly enriched in called peaks. Motifs were identified using HOMER's findMotifsGenome.pl tool. Motifs were called at multiple lengths, and results from all examined lengths are aggregated in the tables.

The following columns are included in the tables:

  •  Motif: logo plot for the identified motif.
  •  IUPAC: sequence of motif in IUPAC code.
  •  Motif length: length of the called motif.
  •  P-value: Significance of the called motif.
  •  -log10(p-value): log transformed significance, a larger number indicates a greater significance.
  •  % Peaks: percent of examined peaks with the identified motif.
  •  % Background: percent of non-peak background regions with the motif.

Please note, this table is interactive. You can search for specific sequences, and can sort the table by any of the columns. The search buttons at thr bottom of the tables allow for specific filtering by sequence or motif length.

ENCSR488JKQ (HepG2) Motifs


No significant motif were identified


ENCSR964VOX (K562) Motifs


Motif IUPAC sequence Motif length P-value -log10(p-value) % Peaks % Background
UCDUCCUUGGAU121e-1536.715.340.02